Eine frisch beringte Nachtigall wird aus der Hand fliegengelassen



Ich forsche darüber, wie sich Tiere (und auch Menschen) mit Lautäußerungen verständigen. Im Fokus steht dabei der Gesang der Nachtigall.

Doch auch für andere Tiere beforschte ich die „vier großen Ws“ des Tierverhalten: wie, warum und „zweimal“ seit wann (im evolutionären sowie im ontogenetischen Kontext) ein bestimmtes Verhalten auftritt. Hier aufgelistet sind die aus dieser Forschung resultierenden wissenschaftliche Studien.


  • Landgraf, C., Wilhelm, K., Wirth, J., Weiss, M. & Kipper, S. (2017). Affairs happen—to whom? A study on extrapair paternity in common nightingales. Current Zoology 2017 zox024. doi: 10.1093/cz/zox024
  • Kipper, S., Sellar, P., & Barlow, C. (2017). The diurnal song of Common nightingales (Luscinia megarhynchos) during the non-breeding period in The Gambia, West-Africa compared to song during the European breeding-season. Journal of Ornithology 158: 223-231. doi:10.1007/s10336-016-1364-0
  • Bartsch, C., Weiss, M., & Kipper, S. (2015). Multiple song features are related to paternal effort in Common nightingales. BMC Evolutionary Biology 15: 115,DOI 10.1186/s12862-015-0390-5
  • Bartsch, C., Hultsch, H., Scharff, C., & Kipper, S. (2015). What is the whistle all about? A study on whistle songs, related male characteristics, and female song preferences in common nightingales. Journal of Ornithology 157: 49-60.DOI 10.1007/s10336-015-1245-y
  • Kipper, S., Kiefer, S., Bartsch, C., Weiss, M. (2015). Female calling? Song responses to conspecific call playbacks in nightingales (Luscinia megarhynchos). Animal Behaviour 100: 60-66.http://dx.doi.org/10.1016/j.anbehav.2014.11.011
  • Kiefer, S., Scharff, C., Hultsch, H., Kipper, S. (2014). Learn it now, sing it later? Field and laboratory studies on song repertoire acquisition and song use in nightingales. Naturwissenschaften 101:955-963, doi:10.1007/s00114-014-1236-5
  • Weiss, M., Hultsch, H., Adam, I., Scharff, C., Kipper, S. (2014). The use of network analysis to study complex animal communication systems: A study on nightingale song. Proceedings of the Royal Society B. doi:10.1098/rspb.2014.0460
  • Bartsch, C., Wenchel, R., Kaiser, A., Kipper, S. (2014). Singing onstage: Female and male Common nightingales eavesdrop on song type matching. Behavioral Ecology and Sociobiology 68, 1163-1171. doi: 10.1007/s00265-014-1727-6
  • Weiss, M., Kiefer, S. & Kipper, S. (2012). Buzzwords’ in females ears? The use of buzz songs in the communication of nightingales (Luscinia megarhynchos). PLoS ONE 7(9): e45057. DOI: 10.1371/journal.pone.0045057
  • Bartsch, C., Weiss, M. & Kipper, S. (2012). The return of the intruder: long-term effects of playbacks from different distances in a territorial songbird. Ethology 118: 876-884.
  • Kiefer, S., Scharff, C. & Kipper, S. (2011). Does age matter in song bird vocal interactions? Results from interactive playback experiments. Frontiers in Zoology 8:29 doi:10.1186/1742-9994-8-29.
  • Kiefer, S., Sommer, C., Scharff, C., & Kipper S. (2010). Singing the popular songs? Nightingales share more song types with their breeding population in their second season than in their first. Ethology 116: 619-626.
  • Kiefer, S., Sommer, C., Scharff, C., Kipper S. & Mundry, R. (2009). Tuning towards tomorrow? Common nightingales Luscinia megarhynchos change and increase their song repertoires from the first to the second breeding season. Journal of Avian Biology 40: 231-236.
  • Kipper, S., Mundry, R., Sommer, C., Hultsch, H. & Todt, D. (2006). Larger nightingales (Luscinia megarhynchos) have larger song repertoires and arrive earlier on their breeding grounds. Animal Behaviour 71:211-217.
  • Kiefer, S., Spiess, A., Kipper, S., Mundry, R., Sommer, C., Hultsch, H. & Todt, D. (2006). First year Common nightingales (Luscinia megarhynchos) have smaller song-repertoire sizes than older males.  Ethology 112: 1217-1224.
  • Naguib, M. & Kipper, S. (2006). Effects of different levels of song overlapping on singing behaviour in male territorial nightingales. Behavioral Ecology and Sociobiology 59: 419-426.
  • Kipper, S., Mundry, R., Hultsch, H. & Todt, D. (2004): Long-term persistence of song performance rules in nightingales (Luscinia megarhynchos): A longitudinal field study on repertoire size and composition. Behaviour 141: 371-390.


  • Fischer, L., Möller Palau-Ribes, F., Kipper, S., Weiss, M., Landgraf, C., & Lierz, M. (2022). Absence of Mycoplasma spp. in nightingales (Luscinia megarhynchos) and blue (Cyanistes caeruleus) and great tits (Parus major) in Germany and its potential implication for evolutionary studies in birds. European Journal of Wildlife Research, 68(1), 1-4
  • Mäntylä, E.,Kipper, S. & Hilker, M. (2020). Insectivorous birds can see and smell systemically herbivore-induced pines. Ecology and Evolution. DOI: 10.1002/ECE3.6622
  • Mäntylä, E., Kleier, S., Lindstedt, C., Kipper, S. & Hilker, M. (2018). Insectivorous Birds Are Attracted by Plant Traits Induced by Insect Egg Deposition. Journal of Chemical Ecology 44: 1127-1138.doi: https://doi.org/10.1007/s10886-018-1034-1
  • Mäntylä, E., Kleier, S., Kipper, S., & Hilker, M. (2017). The attraction of insectivorous birds to herbivore-damaged pines. Journal of Ornithology 158: 479-491.DOI 10.1007/s10336-016-1412-9
  • Honarmand, M., Thompson, C.K.T., Schatton, A., Kipper, S. & Scharff, C. (2015). Early nutritional stress negatively affects neural recruitment to avian song system nucleus HVC. Journal of Developmental Neuroscience DOI 10.1002/dneu.22302
  • Bessert-Nettelbeck, M., Kipper, S., Bartsch, C., Voigt-Heucke, S.L. (2014). Similar, yet different: male Reed Buntings (Emberiza schoeniclus) show high individual differences in song composition, rates of syllable sharing and use. Journal of Ornithology 155: 689-700. doi: 10.1007/s10336-014-1052-x
  • Vokurkova, J., Petruskova, T., Reifova, R., Kozman, A., Morkovsky, L., Kipper, S., Weiss, M., Reif, J., Dolata, P.T., Petrusek, A. (2013). The causes and evolutionary consequences of miexed singing in two hybridizing songbird species (Luscinia spp.) PLoS ONE 8(4): e60172. doi:10.1371/journal.pone.0060172
  • Apfelbeck, B., Mortega, K., Kiefer, S., Kipper, S., Goymann, W. (2013). Life-history and hormonal control of aggression in black redstarts: Blocking testosterone does not decrease territorial aggression, but changes the emphasis of vocal behaviours during simulated territorial intrusions. Frontiers in Zoology,  doi:10.1186/1742-9994-10-8.
  • Apfelbeck, B., Mortega, K., Kiefer, S., Kipper, S., Vellema, M., Villavicencio, C., Gahr, M., Goymann, W. (2013).  Associated and disassociated changes in hormones, song, behavior and brain receptor expression between life-cycle stages in male black redstarts, Phoenicurus ochruros. General & Comparative Endocrinology 184, 93-102.
  • Apfelbeck, B., Kiefer, S., Mortega, K., Goymann, W. & Kipper, S. (2012). Testosterone affects song modulation during simulated territorial intrusions in male black redstarts (Phoenicurus ochruros).  PLoS ONE 7(12): e52009. DOI: 10.1371/journal.pone.0052009
  • Searcy, W. A., Peters, S., Kipper, S., & Nowicki, S. (2010).  Female Sparrows Use Song to Assess Male Developmental History. Behavioural Ecology and Sociobiology 64: 1343-1349.
  • Kipper, S. & Kiefer, S. (2010). Age-related changes in bird’s singing styles: on fresh tunes and fading voices? (Review). Advances in the Study of Behaviour 41: 77-118.


  • Voigt-Heucke, S.L., Zimmer, S., & Kipper, S. (2016). Does Interspecific Eavesdropping Promote Aerial Aggregations in European Pipistrelle Bats During Autumn? Ethology 122: 745–757. doi:10.1111/eth.12519
  • Bendler , C., Kipper, S., Schrader, L. (2014). Vigilance and roosting behaviour of laying hens on different perch heights. Applied Animal Behaviour Science 157:  93-99.doi:10.1016/j.applanim.2014.06.004
  • Brumm, H., Kipper, S., Riechelmann, C. & Todt, D. (2005). Do Barbary macaques ‘comment’ on what they see? A first report on vocalizations accompanying interactions of third parties. Primates 46: 141-144.
  • Kipper, S. & Todt, D. (2003): Dynamic-acoustic variation causes differences in evaluations of laughter. Perceptual and Motor Skills 96: 799-809.
  • Kipper, S. & Todt, D. (2003): The role of rhythm and pitch in the evaluation of human laughter. Journal of Nonverbal Behavior 27 (4): 255-272.
  • Kipper, S. & Todt, D. (2002): The use of vocal signals in the social play of Barbary macaques. Primates 42 (1): 3-17.
  • Kipper, S. & Todt, D. (2001): Variation of sound parameters affects the evaluation of human laughter. Behaviour 138: 1161-1178.
  • Ackermann, T., Kipper, S., Simon, H.J. (2015). “Wenn Bert und Busstop balzen…” – Tiernamen in verhaltensbiologischer Forschung. Beiträge zur Namensforschung, Universitätsverlag Winter, Heidelberg , pp 471-494.
  • Kipper, S. & Todt, D. (2014). Series of similar vocal elements as a Crucial Acoustic Structure in Human Laughter. In Trouvain, J. & Campbell, N.: Phonetics of laughing. Saarbrücken: Universaar – Saarland University Press.
  • Vettin, J. & Kipper, S. (2014). On the role of laughter in conversation: Evaluation differences between short and long laughter bouts. In Trouvain, J. & Campbell, N.: Phonetics of laughing. Saarbrücken: Universaar – Saarland University Press
  • Kipper, S. & Todt, D. (2005). The sound of laughter – recent concepts and findings in research into laughter vocalisations. In: Garfitt, T., McMorran, E. & Taylor, J. (eds.), The Anatomy of Laughter. (Studies in Comparative Literature 8). Legenda, Modern Humanities Research Association and Maney Publishing: London, pp. 24-33.
  • Kipper, S. & Todt, D. (2004). Verhaltensbiologie: Wissenschaft an der Schnittstelle zwischen Tier und Mensch. In: J. Kallinich, & G. Spengler, (Ed.), Tierische Kommunikation.(Kataloge der Museumsstiftung Post und Telekommunikation, Bd.19). Edition Braus: Heidelberg, pp 133-142.
  • Todt, D. & Kipper, S. (2003). Der Begriff der Kommunikation in den Biowissenschaften. In: H. Richter, H. W. Schmitz (Ed.), Kommunikation – ein Schlüsselbegriff der Humanwissenschaften? Nodus Publikationen: Münster, pp. 25-59.